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Details of Dr Ricardo Serrão Serrao Santos   


Gender male
Job title Director
Department Department of Oceanography and Fisheries (DOP)
Universidade dos Açores, Departamento de Oceanografia e Pescas ( DOP/UAç)
Cais de Santa Cruz Horta Açores PT - 9901-862
IMAR Departamento de Oceanografia e Pescas Universidade dos Açores
Tel +351.292200400
Fax +351.292200411
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Website http://www.horta.uac.pt, http://url2
Social Media Links
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Degree PhD
Job type Teaching/Education , Research , Natural Resource Management , Institutional Management
Research Area(s) Biological Oceanography, Marine Ecology , Policy, Law, Economics, Management , Biological Oceanography, Marine Ecology , Policy, Law, Economics, Management
Research Region 36, 70
PUBLICATIONS (as reported by Dr Ricardo Serrão SANTOS)

Walton, M. J., M. A. Silva, S. M. Magalhães, R. Prieto & R. S. Santos (2007). Using blubber biopsies to provide ecological information about bottlenose dolphins Tursiops truncatus around the Azores. Journal of the Marine Biological Association of the U.K., 87: 223–230.
Lobo-da-Cunha, A., E. Kádár & R. S. Santos 2006. Histochemical and ultrastructural characterisation of mantle storage cells in the hydrothermal-vent bivalve Bathymodiolus azoricus. Marine Biology, 150: 253-260.
Ros, A., R. S. Santos & R. F. Oliveira. (2006). Regulation of immunocompetence by different androgen metabolites in a blenny with alternative reproductive tactics. Journal of Experimental Zoology Part A: Comparative Experimental Biology, Volume 305A, 12: 986-994.
Kádár, E., V. Costa & R. S. Santos 2006. Distribution of micro-essential (Fe, Cu, Zn) and toxic (Hg) metals in tissues of two nutritionally distinct hydrothermal shrimps. Science of the Total Environment, 358, (1-3): 143-150
Martins, I., V. Costa, F. Porteiro, A. Colaço & R.S. Santos 2006. Mercury concentrations in fish species caught at Mid-Atlantic Ridge hydrothermal vent fields. Marine Ecology - Progress Series, 320: 253-258
Cardigos, F., F. Tempera, S. Ávila, J. Gonçalves, A. Colaço & R.S. Santos 2006. Non-indigenous marine species of the Azores. Helgoland Marine Research, 60: 160-169.
Cardigos, F., A. Colaço, P. R. Dando, S. P. Ávila, P. M. Sarradin, F. Tempera, F., P. Conceição, A. Pascoal & R. S. Santos 2005. Shallow water hydrothermal vent field fluids and communities of the D. João de Castro Seamount (Azores). Chemical Geology, 224 (1-3): 153-168.
Barreiros, J. P., V. Figna, M. Hostim-Silva & R. S. Santos 2005. Diel seasonality of a shallow-water fish assemblage in a sandy beach at Canto Grande, Santa Catarina, Brazil. Journal of Coastal Research, Sp. Iss. 42: 343-347.
Kádár, E., V. Costa, I. Martins, R. S. Santos & J. J. Powell 2005. Enrichment in trace metals (Al, Mn, Co, Cu, Mo, Cd, Fe, Zn, Pb and Hg) of macro-invertebrate habitats at hydrothermal vents along the Mid Atlantic Ridge. Hydrobiologia, 548: 191-205.
Figueiredo, M., T. Morato, J. P. Barreiros, P. Afonso & R. S. Santos 2005. Feeding ecology of the white seabream, Diplodus sargus, and the ballan wrasse, Labrus bergylta, in the Azores. Fisheries Research, 75: 107–119.
Oliveira, R. F., N. Carvalho, J. A. Miranda, E. J. Gonçalves, M. S. Grober & R. S. Santos 2002. The relationship between the presence of satellite males and nest-holders' mating success in the Azorean rock-pool blenny, Parablennius sanguinolentus parvicornis. Ethology 108: 223-235.
Monteiro, L. R., V. Costa, R. W. Furness & R. S. Santos 1996. Mercury concentrations in prey fish indicate enhanced bioaccumulation in mesopelagic environments. Marine Ecology Progress Series, 141: 21-25.
Almada, V. C. & R. S. Santos 1995. Parental care in the rocky intertidal: a case study of adaptation and exaptation in Mediterranean and Atlantic blennies. Reviews of Fish Biology and Fisheries, 5: 23-37.
Santos, R. S. 1995. Anatomy and histology of secondary sexual characters, gonads and liver of the rock-pool blenny (Parablennius sanguinolentus parvicornis) (Pisces: Blenniidae) of the Azores. Arquipélago. Life and Marine Sciences, 13A: 21-38.
Nash, R. D. M., R. S. Santos, A. J. Geffen, G. Hughes & T. R. Ellis 1994. Diel variability in catch rate of juvenile flatfish on two small nursery grounds (Port Erin Bay, Isle of Man and Porto Pim Bay, Faial, Azores). Journal of Fish Biology, 44: 35-45.
Santos, R. S. & J. P. Barreiros 1993. The ethogram of Parablennius sanguinolentus parvicornis (Pisces: Blenniidae) of the Azores. Arquipélago. Life and Marine Sciences 11A: 73-90.
Nash, R. D. M., A. Geffen & R. S. Santos 1991. The wyde-flounder, Bothus podas Delaroche, a singular flat fish in varied shallow-water habitats of the Azores. Netherlands Journal of Sea Research, 27(3/4): 367-373.
Santos, R. S., V. Almada & A. J. F. Santos 1989. Field experiments and observations on homing and territoriality in intertidal fishes. Pp. 623-632. In: R. J. Blanchard, P. F. Brain, D. C. Blanchard & S. Parmigiani (Eds.) EthoExperimental Analysis of Behaviour, Kluwer Academic Press, Dordrecht.
Santos, R. S. & V. Almada 1988. Intraspecific variations in reproductive tactics in males of the rocky intertidal fish Blennius sanguinolentus in Azores. Pp. 421-447. In: G. Chellazzi & M. Vannini (Eds.): Behavioral Adaptation to Intertidal Life. Plenum Press, New York.
Ros A. F. H, N. Bouton, R. S. Santos & R. F. Oliveira 2006. Alternative male reproductive tactics and the immunocompetence handicap in the Azorean rock pool blenny, Parablennius parvicornis. Proceedings of the Royal Society B: Biological Sciences, 273 (1589), 901–909.
Colaço, A., P. Bustamante, Y. Fouquet, P.-M. Sarradin & R. S. Santos 2006. Bioaccumulation of Hg, Cu, and Zn in the Azores Triple Junction hydrothermal vent field food web. Chemosphere 65 (11): 2260–2267.
Dixon, D. R., D. M. Lowe, P. I. Miller, G. Villemin, A. Colaço, R. S. Santos, L. R. J. Dixon 2006. Evidence for seasonal reproduction in the Atlantic vent mussel Bathymodiolus azoricus, and an apparent link to the timing of photosynthetic primary Journal of the Marine Biological Association of the UK, 86 (6): 1363-1371.
Martins, I., V. Costa, F. Porteiro & R.S. Santos 2006. Temporal and spatial changes in mercury concentrations in the North Atlantic as indicated by museum specimens of glacier lanternfish Benthosema glaciale (Pisces: Myctophidae). Environmental Toxicology, 21 (5): 528-532.
Santos, R. S., S. J. Hawkins & R. D. M. Nash 1996. Reproductive phenology of the Azorean rock-pool blenny, a fish with alternative mating tactics. Journal of Fish Biology, 48 (5): 842-858.
Santos, R. S. & R. D. M. Nash 1995. Seasonal changes in a sandy beach fish assemblage at Porto Pim, Faial, Azores. Estuarine, Coastal and Shelf Science, 41 (5): 579-591.
Santos, R. S., S. Hawkins, L. R. Monteiro, M. Alves & E. J. Isidro 1995. Marine research, resources and conservation in the Azores. Aquatic Conservation: Marine and Freshwater Ecosystems, 5 (4): 311-354.
Almada, F., V. C. Almada, V. Domingues, A. Brito & R. S. Santos 2005. Molecular validation of the specific status of Parablennius sanguinolentus and Parablennius parvicornis (Pisces: Blenniidae). Scientia Marina, 69 (4): 519-523.
Colaço, A., C. Raghukumar, C. Mohandass, F. Cardigos & R. S. Santos 2006. Effect of shallow-water venting in Azores on a few marine biota. Cahiers de Biologie Marine, 47 (4): 359-364.
Domingues, V. S., V. C. Almada, R. S. Santos, A. Brito & G. Bernardi 2007. Phylogeography and evolution of the triplefin Tripterygion delaisi (Pisces, Blennioidei). Marine Biology, 150 (3): 509-519.
Kádár, E. V., Costa, V., R.S. Santos & H. Lopes 2005. Behavioural response to the bioavailability of inorganic hg in the hydrothermal mussel Bathymodiolus azoricus. Journal of Experimental Biology, 208 (3): 505-513.
Santos, R. S., R. D. M. Nash & S. J. Hawkins 1995. Age, growth and sex ratio of the Azorean rock-pool blenny, Parablennius sanguinolentus parvicornis. Journal of the Marine Biological Association of the United Kingdom, 75 (3): 751-754.
Patzner, R. A. & R. S. Santos 1992. Field observations on the association between the clingfish Diplecogaster bimaculata pectoralis Briggs 1955 and different species of sea urchins at the Azores. Zeitschrift für Fischkunde, 1 (2):157-161.
Spence, S., S. J. Hawkins & R. S. Santos 1990. Thais haemastoma: An exhibitor of "imposex" and potential biological indicator of Tributyltin. P.S.Z.N.I. - Marine Ecology, 11(2): 147-156.
Nash, R. D. M. & R. S. Santos 1993. The occurrence of the lesser weever (Echiichthys vipera) (Cuvier 1829) in the Azores. Journal of Fish Biology, 43 (2): 317-319.
De Girolamo, M. de, A. Grapputo, L. Zane, R . S. Santos & A. Pallavicini 2005. Isolation of seven polymorphic microsatellites in Ophioblennius atlanticus atlanticus (Perciformes, Blenniidae). Molecular Ecology Notes 5 (2): 334-336.
Colaço A., I. Martins, M. Laranjo, L. Pires, C. Leal, C. Prieto, V. Costa, H. Lopes, D. Rosa, P. R. Dando & R. S. Santos 2006. Annual spawning of the hydrothermal vent mussel, Bathymodiolus azoricus, under controlled aquarium conditions at atmospheric pressure. Journal of Experimental Marine Biology and Ecology, 333 (2): 166-171.
Santos, R.S., J. Gonçalves & E. Isidro 1995. Marine Research: The role of the Department of Oceanography and Fisheries of the University of the Azores. Higher Education Policy, 8 (2): 25-28.
Kádár, E., R. S. Santos & J. J. Powell 2006. Biological factors influencing tissue compartmentalization of trace metals in the deep-sea hydrothermal vent bivalve Bathymodiolus azoricus at geochemically distinct vent sites of the Mid-Atlantic Ridge. Environmental Research, 101 (2): 221-229.
Santos, R. S. 1995. Allopaternal care in the redlip blenny. Journal of Fish Biology, 47 (2): 350-353.
Kádár, E., R. Bettencourt, V. Costa, R. S. Santos, A. Lobo-da-Cunha & P. Dando 2005. Experimentally induced endosymbiont loss and re-acquirement in the hydrothermal vent bivalve Bathymodiolus azoricus. Journal of Experimental Marine Biology and Ecology, 318 (1): 99-110.
Stefanni, S., V. Domingues, N. Bouton, V. Almada & R. S. Santos 2006. Phylogeny of the shanny, Lipophrys pholis, from the NE Atlantic using mitochondrial DNA markers. Molecular Phylogenetics and Evolution, 39 (1): 282-287.
Kádár, E., A. Lobo-Da-Cunha, R. S. Santos & P. Dando 2006. Spermatogenesis of Bathymodiolus azoricus in captivity matching reproductive behaviour at deep-sea hydrothermal vents. Journal of Experimental Marine Biology and Ecology, 335 (1): 19-26
Ros A.F.H., N. Bouton, R.S. Santos & R. F. Oliveira 2004. Are aggressive fathers immunosuppressed? Alternative reproductive tactics, androgens and immunocompetence in the Azorean rock-pool blenny. Hormones and Behavior 46 (1): 120-120.
Afonso, P. & R. S. Santos 2005. Within-nest spawning site preferences of female bluefin damselfish: the effect of early-stage eggs. Acta Ethologica, 8 (1): 5-11
os A. F. H, C. Ferreira, R. S. Santos & R. F. Oliveira 2005 An experimental test of the immunocompetence handicap hypothesis in the Azorean rock-pool blenny. Hormones and Behavior 48 (1): 123-123.
Santos, R. S. & R. D. M. Nash 1996. Seasonal variations in injuries suffered by individuals of the Azorean rock-pool blenny, Parablennius sanguinolentus parvicornis. Copeia, 1996 (1): 216-219.
Kádár, E., V. Costa, R. S. Santos & J. J. Powell 2006. Tissue partitioning of micro-essential metals in the vent bivalve Bathymodiolus azoricus and associated organisms (endosymbiont bacteria and parasite polychaete) from geochemically distinct vents of the Mid-Atlantic Ridge. Journal of Sea Research, 56 (1): 45-52.
Domingues, V. S., R. S. Santos, A. Brito & V. C. Almada 2006. Historical population dynamics and demography of the eastern Atlantic pomacentrid Chromis limbata (Valenciennes, 1833). Molecular Phylogenetics and Evolution, 40 (1): 139-147.

Group(s): IOC , Ocean Sciences
Created: 2004-11-08 Last Updated: 2011-03-25 by Luciano FONSECA